Police are appealing for information and witnesses following the collision in Haywards Heath.
Emergency services were called to the southbound carriageway of the A23, between Hickstead and the A272 junction, just after 10.40am on Sunday.
The collision involved a white Peugeot and a Triumph motorcycle.
The rider of the motorcycle, a 65-year-old man from Brighton, sustained serious injuries and was taken to hospital.
A Sussex Police spokesman said: “Officers are appealing for information and want to speak to anyone who saw, or who has dashcam footage, of what happened.
“Information can be reported to police online or via 101, quoting serial 404 of 9/3.”
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