One is at the Seven Dials Roundabout in Dyke Road, Brighton, and the other in High Street, Lewes.
They will be the fourth and fifth branches in Sussex after a third opened in Horsham last summer.
Brighton’s New Road Gail’s and the branch in Church Road, Hove, have both proved popular.
The new one at Seven Dials takes the place of pop-up wine bar Night Shift which filled the space temporarily after the Small Batch Coffee Roasters closed last March.
Lewes’s first branch opens in what was the Natwest bank on the corner of Friars Walk. The site has been empty since January last year.
As well as breads and pastries, the cafés will serve sandwiches, cakes and coffee.
Gail’s Seven Dials will open its doors on Wednesday, February 28, and the Lewes branch will open the following day.
News of the chain’s latest Brighton opening drew a mixed response when branding was plastered on the store in January.
Writing on Facebook, Nick Saxon said: “There are already plenty of cafes, and sandwich shops in the Dials.
“This will probably result in a few of the local independents closing. They will employ locals but at the expense of other places closing.
“All of their products are made off site. It’s overpriced in my opinion.”
The word “boring” and a large penis were spray-painted on the building last month.
A spokesman for Gail’s said the response was a shame.
“It is sad to see comments from a small proportion of the community in Seven Dials and the recent graffiti outside our bakery,” he said.
“We’ve been serving Brighton craft baking and speciality coffee for over seven years now and really love this neighbourhood.”
Lynne Knight, who founded the homeless charity Knight Support, said: “They donate their end of day produce that isn’t sold to our homeless and other supported accommodation places for people suffering hardship and poverty free of charge.
“They went out of their way to make this happen to help people in need and to save good food being thrown away.”
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