The sale, which goes on until January 29, sees the price of selected fares slashed by up to half.
Advance single tickets, which allow passengers to travel on specific trains only, are part of the discount.
For journeys between January 30 and March 15, an advance single ticket for the journey from Brighton to London Victoria could cost as little as £3.
The deal is also on for select tickets from Lewes to London Victoria.
The sale, introduced by the government, will boost tourism and encourage sustainable travel, it said.
Transport Secretary Mark Harper said: “The return of the Great British Rail Sale is good news for passengers, following the success of the previous sale which saw passengers benefit from around £7 million in savings on their travel costs.
“With discounts on more than a million tickets, there’ll be plenty of opportunity to connect with friends and family and explore great destinations across the country – I hope passengers make the most of this sale and choose to travel by rail.”
Jacqueline Starr, chief executive of the rail delivery group said: “Train travel is the greenest way to explore our towns and cities as well as connect with your family and friends because nothing beats being there in person for those special moments.
“Following the success of the Great British Rail Sale in 2022, we are really excited that it’s coming back.”
The Great British Railways Transition Team (GBRTT) estimate that the first Great British Rail Sale in 2022 saw passengers save around £7 million on rail tickets and encouraged around 70,000 adults who hadn’t travelled by train since the pandemic to take a trip.
Suzanne Donnelly, passenger revenue director at GBRTT, said: “I’m delighted the rail industry has, once again, come together to deliver another huge round of savings on tickets for passengers.
Only a limited number of tickets are available, and no further sale tickets will go on offer once sold out.
Tickets must be purchased at least seven days in advance.
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